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One more kind of secondary sexual character must here be mentioned: the odour which emanates from so many animals at the breeding season. It is possible that this odour also served at first merely to give notice of the presence of individuals of the other sex, but it soon became an excitant, and as the individuals which caused the greatest degree of excitement were preferred, it reached as high a pitch of perfection as was possible to it. I shall confine myself here to the comparatively recently discovered fragrance of butterflies. Since Fritz Mueller found out that certain Brazilian butterflies gave off fragrance "like a flower," we have become acquainted with many such cases, and we now know that in all lands, not only many diurnal Lepidoptera but nocturnal ones also give off a delicate odour, which is agreeable even to man. The ethereal oil to which this fragrance is due is secreted by the skin-cells, usually of the wing, as I showed soon after the discovery of the _scent-scales_. This is the case in the males; the females have no _special_ scent-scales recognisable as such by their form, but they must, nevertheless, give off an extremely delicate fragrance, although our imperfect organ of smell cannot perceive it, for the males become aware of the presence of a female, even at night, from a long distance off, and gather round her. We may therefore conclude, that both sexes have long given forth a very delicate perfume, which announced their presence to others of the same species, and that in many species (_not in all_) these small beginnings become, in the males, particularly strong scent-scales of characteristic form (lute, brush, or lyre-shaped).

The cruelty of the Patrician creditors was the most pressing evil, and led to the first reform. In B.C. 494 the Plebeians, after a campaign against the Volscians, instead of returning to Rome, retired to the Sacred Mount, a hill about two miles from the city, near the junction of the Arno and the Tiber. Here they determined to settle and found a new town, leaving Rome to the Patricians and their clients. This event is known as the _Secession to the Sacred Mount_. The Patricians, alarmed, sent several of their number to persuade the Plebeians to return. Among the deputies was the aged Menenius Agrippa, who had great influence with the Plebeians. He related to them the celebrated fable of the Belly and the Members.

For a long time the marvellously complex _radiate_ and _lattice-work_ skeletons of Radiolarians were regarded as a mere outflow of "Nature's infinite wealth of form," as an instance of a purely morphological character with no biological significance. But recent investigations have shown that these, too, have an adaptive significance (Haecker). The same thing has been shown by Schuett in regard to the lowly unicellular plants, the Peridineae, which abound alike on the surface of the ocean and in its depths. It has been shown that the long skeletal processes which grow out from these organisms have significance not merely as a supporting skeleton, but also as an extension of the superficial area, which increases the contact with the water-particles, and prevents the floating organisms from sinking. It has been established that the processes are considerably shorter in the colder layers of the ocean, and that they may be twelve times as long[36] in the warmer layers, thus corresponding to the greater or smaller amount of friction which takes place in the denser and less dense layers of the water.

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